[1] PARK J H, BOLAN N, MEGHARAJ M, et al. Isolation of phosphate solubilizing bacteria and their potential for lead immobilization in soil [J]. Journal of Hazardous Materials, 2011, 185(2/3): 829-836.
[2] TAURIAN T, ANZUAY M S, ANGELINI J G, et al. Phosphate-solubilizing peanut associated bacteria: Screening for plant growth-promoting activities [J]. Plant and Soil, 2010, 329(1/2): 421-431.
[3] BOLAN N S, ADRIANO D C, NAIDU R. Role of phosphorus in (Im)mobilization and bioavailability of heavy metals in the soil-plant system [J]. Reviews of Environmental Contamination and Toxicology, 2003, 177: 1-44.
[4] SINGH B K. Organophosphorus-degrading bacteria: Ecology and industrial applications [J]. Nature Reviews Microbiology, 2009, 7(2): 156-164. doi: 10.1038/nrmicro2050
[5] SON H J, PARK G T, CHA M S, et al. Solubilization of insoluble inorganic phosphates by a novel salt- and pH-tolerant Pantoea agglomerans R-42 isolated from soybean rhizosphere [J]. Bioresource Technology, 2006, 97(2): 204-210. doi: 10.1016/j.biortech.2005.02.021
[6] GILES C D, HSU P C, RICHARDSON A E, et al. Plant assimilation of phosphorus from an insoluble organic form is improved by addition of an organic anion producing Pseudomonas sp. [J]. Soil Biology and Biochemistry, 2014, 68: 263-269. doi: 10.1016/j.soilbio.2013.09.026
[7] HARIPRASAD P, NIRANJANA S R. Isolation and characterization of phosphate solubilizing rhizobacteria to improve plant health of tomato [J]. Plant and Soil, 2009, 316(1/2): 13-24.
[8] RICHARDSON A E, HADOBAS P A, HAYES J E. Extracellular secretion of Aspergillus phytase from Arabidopsis roots enables plants to obtain phosphorus from phytate [J]. The Plant Journal: for Cell and Molecular Biology, 2001, 25(6): 641-649. doi: 10.1046/j.1365-313x.2001.00998.x
[9] NELSON T S, SHIEH T R, WODZINSKI R J, et al. Effect of supplemental phytase on the utilization of phytate phosphorus by chicks [J]. The Journal of Nutrition, 1971, 101(10): 1289-1293. doi: 10.1093/jn/101.10.1289
[10] MANSOTRA P, SHARMA P, SHARMA S. Bioaugmentation of Mesorhizobium Cicer, Pseudomonas spp. and piriformospora indica for sustainable chickpea production [J]. Physiology and Molecular Biology of Plants, 2015, 21(3): 385-393. doi: 10.1007/s12298-015-0296-0
[11] RICHARDSON A E, SIMPSON R J. Soil microorganisms mediating phosphorus availability update on microbial phosphorus [J]. Plant Physiology, 2011, 156(3): 989-996. doi: 10.1104/pp.111.175448
[12] CREA F, de STEFANO C, MILEA D, et al. Formation and stability of phytate complexes in solution [J]. Coordination Chemistry Reviews, 2008, 252(10/11): 1108-1120.
[13] MULLANEY E J, DALY C B, ULLAH A H. Advances in phytase research [J]. Advances in Applied Microbiology, 2000, 47: 157-199.
[14] KUMAR V, SINHA A K, MAKKAR H P S, et al. Dietary roles of phytate and phytase in human nutrition: A review [J]. Food Chemistry, 2010, 120(4): 945-959. doi: 10.1016/j.foodchem.2009.11.052
[15] FEKRI A, TORBATI M, YARI KHOSROWSHAHI A, et al. Functional effects of phytate-degrading, probiotic lactic acid bacteria and yeast strains isolated from Iranian traditional sourdough on the technological and nutritional properties of whole wheat bread [J]. Food Chemistry, 2020, 306: 125620. doi: 10.1016/j.foodchem.2019.125620
[16] SONG H Y, EL SHEIKHA A F, HU D M. The positive impacts of microbial phytase on its nutritional applications [J]. Trends in Food Science and Technology, 2019, 86: 553-562. doi: 10.1016/j.jpgs.2018.12.001
[17] JORQUERA M, MARTÍNEZ O, MARUYAMA F, et al. Current and future biotechnological applications of bacterial phytases and phytase-producing bacteria [J]. Microbes and Environments, 2008, 23(3): 182-191. doi: 10.1264/jsme2.23.182
[18] PANDEY A, SZAKACS G, SOCCOL C R, et al. Production, purification and properties of microbial phytases [J]. Bioresource Technology, 2001, 77(3): 203-214. doi: 10.1016/S0960-8524(00)00139-5
[19] GOLOVAN S, WANG G, ZHANG J, et al. Characterization and overproduction of the Escherichia coli appA encoded bifunctional enzyme that exhibits both phytase and acid phosphatase activities [J]. Canadian Journal of Microbiology, 2000, 46(1): 59-71. doi: 10.1139/cjm-46-1-59
[20] PHILLIPPY B Q, MULLANEY E J. Expression of an Aspergillus niger phytase (phyA) in Escherichia coli [J]. Journal of Agricultural and Food Chemistry, 1997, 45(8): 3337-3342. doi: 10.1021/jf970276z
[21] MULLANEY E J, ULLAH A H J. The term phytase comprises several different classes of enzymes [J]. Biochemical and Biophysical Research Communications, 2003, 312(1): 179-184. doi: 10.1016/j.bbrc.2003.09.176
[22] SHIVANNA G B, GOVINDARAJULU V. Screening of asporogenic mutants of phytase-producing aspergillusniger CFR 335 strain [J]. Microbial Ecology in Health and Disease, 2009, 21(1): 57-63. doi: 10.1080/08910600902745750
[23] MITCHELL D B, VOGEL K, WEIMANN B J, et al. The phytase subfamily of histidine acid phosphatases: Isolation of genes for two novel phytases from the fungi Aspergillus terreus and Myceliophthora thermophila[J]. Microbiology, 1997, 143 (1): 245-252.
[24] OLSTORPE M, SCHNÜRER J, PASSOTH V. Screening of yeast strains for phytase activity [J]. FEMS Yeast Research, 2009, 9(3): 478-488. doi: 10.1111/j.1567-1364.2009.00493.x
[25] NAKAMURA Y, FUKUHARA H, SANO K. Secreted phytase activities of yeasts [J]. Bioscience, Biotechnology, and Biochemistry, 2000, 64(4): 841-844. doi: 10.1271/bbb.64.841
[26] VOHRA A, SATYANARAYANA T. Phytases: microbial sources, production, purification, and potential biotechnological applications [J]. Critical Reviews in Biotechnology, 2003, 23(1): 29-60. doi: 10.1080/713609297
[27] RICHARDSON A E, HADOBAS P A. Soil isolates of Pseudomonas spp. that utilize inositol phosphates [J]. Canadian Journal of Microbiology, 1997, 43(6): 509-516. doi: 10.1139/m97-073
[28] VOLFOVÁ O, DVOŘÁKOVÁ J, HANZLÍKOVÁ A, et al. Phytase from Aspergillus niger [J]. Folia Microbiologica, 1994, 39(6): 481-484. doi: 10.1007/BF02814066
[29] GARGOVA S, ROSHKOVA Z, VANCHEVA G. Screening of fungi for phytase production [J]. Biotechnology Techniques, 1997, 11(4): 221-224. doi: 10.1023/A:1018426119073
[30] LAMBRECHTS C, BOZE H, MOULIN G, et al. Utilization of phytate by some yeasts [J]. Biotechnology Letters, 1992, 14(1): 61-66. doi: 10.1007/BF01030915
[31] SANO K, FUKUHARA H, NAKAMURA Y. Phytase of the yeast Arxula adeninivorans [J]. Biotechnology Letters, 1999, 21(1): 33-38. doi: 10.1023/A:1005438121763
[32] BAE H D, YANKE L J, CHENG K J, et al. A novel staining method for detecting phytase activity [J]. Journal of Microbiological Methods, 1999, 39(1): 17-22. doi: 10.1016/S0167-7012(99)00096-2
[33] ENGELEN A J, van der HEEFT F C, RANDSDORP P H G, et al. Simple and rapid determination of phytase activity [J]. Journal of AOAC INTERNATIONAL, 2020, 77(3): 760-764.
[34] VATS P, BANERJEE U C. Production studies and catalytic properties of phytases (myo-inositol hexakisphosphate phosphohydrolases): An overview [J]. Enzyme and Microbial Technology, 2004, 35(1): 3-14. doi: 10.1016/j.enzmictec.2004.03.010
[35] PANDEY A, SOCCOL C R. Bioconversion of biomass: A case study of ligno-cellulosics bioconversions in solid state fermentation [J]. Brazilian Archives of Biology and Technology, 1998, 41(4): 379-390. doi: 10.1590/S1516-89131998000400001
[36] PANDEY A, SOCCOL C R, MITCHELL D. New developments in solid state fermentation: I-bioprocesses and products [J]. Process Biochemistry, 2000, 35(10): 1153-1169. doi: 10.1016/S0032-9592(00)00152-7
[37] PANDEY A, SOCCOL C. Economic utilization of crop residues for value addition: A futuristic approach [J]. Journal of Scientific and Industrial Research, 2000, 59(1): 12-22.
[38] HAN Y W, GALLAGHER D J, WILFRED A G. Phytase production by Aspergillus ficuum on semisolid substrate [J]. Journal of Industrial Microbiology, 1987, 2(4): 195-200. doi: 10.1007/BF01569540
[39] PAPAGIANNI M, NOKES S E, FILER K. Production of phytase by Aspergillus niger in submerged and solid-state fermentation [J]. Process Biochemistry, 1999, 35(3/4): 397-402.
[40] SAJIDAN A, FAROUK A, GREINER R, et al. Molecular and physiological characterisation of a 3-phytase from soil bacterium Klebsiella sp. ASR1 [J]. Applied Microbiology and Biotechnology, 2004, 65(1): 110-118.
[41] SREERAMULU G, SRINIVASA D S, NAND K, et al. Lactobacillus amylovorus as a phytase producer in submerged culture [J]. Letters in Applied Microbiology, 1996, 23(6): 385-388. doi: 10.1111/j.1472-765X.1996.tb01342.x
[42] YANKE L J, BAE H D, SELINGER L B, et al. Phytase activity of anaerobic ruminal bacteria[J]. Microbiology , 1998, 144 ( 6): 1565-1573.
[43] LAN G Q, ABDULLAH N, JALALUDIN S, et al. Culture conditions influencing phytase production of Mitsuokella jalaludinii, a new bacterial species from the rumen of cattle [J]. Journal of Applied Microbiology, 2002, 93(4): 668-674. doi: 10.1046/j.1365-2672.2002.01727.x
[44] HERTER T, BEREZINA O V, ZININ N V, et al. Glucose-1-phosphatase (AgpE) from Enterobacter cloacae displays enhanced phytase activity [J]. Applied Microbiology and Biotechnology, 2006, 70(1): 60-64. doi: 10.1007/s00253-005-0024-8
[45] YOON S J, CHOI Y J, MIN H K, et al. Isolation and identification of phytase-producing bacterium, Enterobacter sp. 4, and enzymatic properties of phytase enzyme [J]. Enzyme and Microbial Technology, 1996, 18(6): 449-454. doi: 10.1016/0141-0229(95)00131-X
[46] TAMBE S M, KAKLIJ G S, KELKAR S M, et al. Two distinct molecular forms of phytase from Klebsiella aerogenes: Evidence for unusually small active enzyme peptide [J]. Journal of Fermentation and Bioengineering, 1994, 77(1): 23-27. doi: 10.1016/0922-338X(94)90202-X
[47] WANG X Y, UPATHAM S, PANBANGRED W, et al. Purification, characterization, gene cloning and sequence analysis of a phytase from Klebsiella pneumoniae subsp. pneumoniae XY-5 [J]. ScienceAsia, 2004, 30: 383-390. doi: 10.2306/scienceasia1513-1874.2004.30.383
[48] GREINER R, HALLER E, KONIETZNY U, et al. Purification and characterization of a phytase from Klebsiella terrigena [J]. Archives of Biochemistry and Biophysics, 1997, 341(2): 201-206. doi: 10.1006/abbi.1997.9942
[49] JAREONKITMONGKOL S, OHYA M, WATANABE R, et al. Partial purification of phytase from a soil isolate bacterium, Klebsiella oxytoca MO-3 [J]. Journal of Fermentation and Bioengineering, 1997, 83(4): 393-394. doi: 10.1016/S0922-338X(97)80149-3
[50] ZININ N V, SERKINA A V, GELFAND M S, et al. Gene cloning, expression and characterization of novel phytase from Obesumbacterium proteus [J]. FEMS Microbiology Letters, 2004, 236(2): 283-290. doi: 10.1111/j.1574-6968.2004.tb09659.x
[51] GREINER R. Purification and properties of a phytate-degrading enzyme from Pantoea agglomerans [J]. The Protein Journal, 2004, 23(8): 567-576. doi: 10.1007/s10930-004-7883-1
[52] VASHISHTH A, RAM S, BENIWAL V. Cereal phytases and their importance in improvement of micronutrients bioavailability [J]. 3 Biotech, 2017, 7(1): 42. doi: 10.1007/s13205-017-0698-5
[53] CHO J, LEE C, KANG S, et al. Molecular cloning of a phytase gene (phy M) from Pseudomonas syringae MOK1 [J]. Current Microbiology, 2005, 51(1): 11-15. doi: 10.1007/s00284-005-4482-0
[54] IN M J, JANG E S, KIM Y, et al. Purification and properties of an extracellular acid phytase from Pseudomonas fragi Y9451 [J]. Journal of Microbiology and Biotechnology, 2004, 14(5): 1004-1008.
[55] HUANG H Q, LUO H Y, YANG P L, et al. A novel phytase with preferable characteristics from Yersinia intermedia [J]. Biochemical and Biophysical Research Communications, 2006, 350(4): 884-889. doi: 10.1016/j.bbrc.2006.09.118
[56] GEORGE T S, SIMPSON R J, GREGORY P J, et al. Differential interaction of Aspergillus niger and Peniophora lycii phytases with soil particles affects the hydrolysis of inositol phosphates [J]. Soil Biology and Biochemistry, 2007, 39(3): 793-803. doi: 10.1016/j.soilbio.2006.09.029
[57] SHIEH T R, WARE J H. Survey of microorganism for the production of extracellular phytase [J]. Applied Microbiology, 1968, 16(9): 1348-1351. doi: 10.1128/am.16.9.1348-1351.1968
[58] VATS P, BANERJEE U C. Biochemical characterisation of extracellular phytase (myo-inositol hexakisphosphate phosphohydrolase) from a hyper-producing strain of Aspergillus niger van Teighem [J]. Journal of Industrial Microbiology and Biotechnology, 2005, 32(4): 141-147. doi: 10.1007/s10295-005-0214-5
[59] HA N C, KIM Y O, OH T K, et al. Preliminary X-ray crystallographic analysis of a novel phytase from a strain [J]. Acta Crystallographica Section D Biological Crystallography, 1999, 55(3): 691-693. doi: 10.1107/S0907444998015285
[60] TYE A, SIU F, LEUNG T, et al. Molecular cloning and the biochemical characterization of two novel phytases from B. subtilis168 and B. licheniformis [J]. Applied Microbiology and Biotechnology, 2002, 59(2/3): 190-197.
[61] KEROVUO J, LAURAEUS M, NURMINEN P, et al. Isolation, characterization, molecular gene cloning, and sequencing of a novel phytase from Bacillus subtilis [J]. Applied and Environmental Microbiology, 1998, 64(6): 2079-2085. doi: 10.1128/AEM.64.6.2079-2085.1998
[62] KIM Y O, LEE J K, KIM H K, et al. Cloning of the thermostable phytase gene (phy) from Bacillus sp. DS11 and its overexpression in Escherichia coli [J]. FEMS Microbiology Letters, 1998, 162(1): 185-191. doi: 10.1111/j.1574-6968.1998.tb12997.x
[63] KIM H W, KIM Y O, LEE J H, et al. Isolation and characterization of a phytase with improved properties from Citrobacter braakii [J]. Biotechnology Letters, 2003, 25(15): 1231-1234. doi: 10.1023/A:1025020309596
[64] PAYNTER M J, ELSDEN S R. Mechanism of propionate formation by Selenomonas ruminantium, a rumen micro-organism [J]. Journal of General Microbiology, 1970, 61(1): 1-7. doi: 10.1099/00221287-61-1-1
[65] IRVING G, COSGROVE D J. Inositol phosphate phosphatases of microbiological origin. some properties of a partially purified bacterial (Pseudomonas sp. ) phytase [J]. Australian Journal of Biological Sciences, 1971, 24(3): 547. doi: 10.1071/BI9710547
[66] LI R J, LU W J, GUO C J, et al. Molecular characterization and functional analysis of OsPHY1, a purple acid phosphatase (PAP)-type phytase gene in rice (Oryza sativa L. ) [J]. Journal of Integrative Agriculture, 2012, 11(8): 1217-1226. doi: 10.1016/S2095-3119(12)60118-X
[67] IDRISS E E, MAKAREWICZ O, FAROUK A, et al. Extracellular phytase activity of Bacillus amyloliquefaciens FZB45 contributes to its plant-growth-promoting effect[J]. Microbiology , 2002, 148(7): 2097-2109.
[68] POWAR V K, JAGANNATHAN V. Purification and properties of phytate-specific phosphatase from Bacillus subtilis [J]. Journal of Bacteriology, 1982, 151(3): 1102-1108. doi: 10.1128/jb.151.3.1102-1108.1982
[69] KUMAR V, SANGWAN P, VERMA A K, et al. Molecular and biochemical characteristics of recombinant β-propeller phytase from Bacillus licheniformis strain PB-13 with potential application in aquafeed [J]. Applied Biochemistry and Biotechnology, 2014, 173(2): 646-659. doi: 10.1007/s12010-014-0871-9
[70] CHOI Y M, SUH H J, KIM J M. Purification and properties of extracellular phytase from Bacillus sp. KHU-10 [J]. Journal of Protein Chemistry, 2001, 20(4): 287-292. doi: 10.1023/A:1010945416862
[71] GREINER R, KONIETZNY U, JANY K D. Purification and characterization of two phytases from Escherichia coli [J]. Archives of Biochemistry and Biophysics, 1993, 303(1): 107-113. doi: 10.1006/abbi.1993.1261
[72] GREINER R, CARLSSON N G. Myo-Inositol phosphate isomers generated by the action of a phytate-degrading enzyme from Klebsiella terrigena on phytate [J]. Canadian Journal of Microbiology, 2006, 52(8): 759-768. doi: 10.1139/w06-028
[73] GREINER R. Degradation of myo-inositol hexakisphosphate by a phytate-degrading enzyme from Pantoea agglomerans [J]. The Protein Journal, 2004, 23(8): 577-585. doi: 10.1007/s10930-004-7884-0
[74] GREINER R, FAROUK A, ALMINGER M L, et al. The pathway of dephosphorylation of myo-inositol hexakisphosphate by phytate-degrading enzymes of different Bacillus spp. [J]. Canadian Journal of Microbiology, 2002, 48(11): 986-994. doi: 10.1139/w02-097
[75] PALACIOS M C, HAROS M, ROSELL C M, et al. Characterization of an acid phosphatase from Lactobacillus pentosus: Regulation and biochemical properties [J]. Journal of Applied Microbiology, 2005, 98(1): 229-237. doi: 10.1111/j.1365-2672.2004.02447.x
[76] ZAMUDIO M, GONZÁLEZ A, MEDINA J A. Lactobacillus plantarum phytase activity is due to non-specific acid phosphatase [J]. Letters in Applied Microbiology, 2001, 32(3): 181-184. doi: 10.1046/j.1472-765x.2001.00890.x
[77] YANKE L J, SELINGER L B, CHENG K J. Phytase activity of Selenomonas ruminantium: A preliminary characterization [J]. Letters in Applied Microbiology, 1999, 29(1): 20-25. doi: 10.1046/j.1365-2672.1999.00568.x
[78] SHAH V, PAREKH L J. Phytase from Klebsiella sp. No. PG–2: Purification and properties [J]. Indian Journal of Biochemistry and Biophysics, 1990, 27(2): 98-102.
[79] LASSEN S F, BREINHOLT J, ØSTERGAARD P R, et al. Expression, gene cloning, and characterization of five novel phytases from four basidiomycete fungi: Peniophora lycii,Agrocybe pediades, a Ceriporia sp., and Trametes pubescens [J]. Applied and Environmental Microbiology, 2001, 67(10): 4701-4707. doi: 10.1128/AEM.67.10.4701-4707.2001
[80] SHANKS R M Q, DASHIFF A, ALSTER J S, et al. Isolation and identification of a bacteriocin with antibacterial and antibiofilm activity from Citrobacter freundii [J]. Archives of Microbiology, 2012, 194(7): 575-587. doi: 10.1007/s00203-012-0793-2
[81] LI C, LIN Y, ZHENG X Y, et al. Combined strategies for improving expression of Citrobacter amalonaticus phytase in Pichia pastoris [J]. BMC Biotechnology, 2015, 15: 88. doi: 10.1186/s12896-015-0204-2
[82] BERKA R M, REY M W, BROWN K M, et al. Molecular characterization and expression of a phytase gene from the thermophilic fungus Thermomyces lanuginosus [J]. Applied and Environmental Microbiology, 1998, 64(11): 4423-4427. doi: 10.1128/AEM.64.11.4423-4427.1998
[83] WYSS M, PASAMONTES L, FRIEDLEIN A, et al. Biophysical characterization of fungal phytases (myo-inositol hexakisphosphate phosphohydrolases): Molecular size, glycosylation pattern, and engineering of proteolytic resistance [J]. Applied and Environmental Microbiology, 1999, 65(2): 359-366. doi: 10.1128/AEM.65.2.359-366.1999
[84] FU S J, SUN J Y, QIAN L C, et al. Bacillus phytases: Present scenario and future perspectives [J]. Applied Biochemistry and Biotechnology, 2008, 151(1): 1-8. doi: 10.1007/s12010-008-8158-7
[85] GU W N, HUANG H Q, MENG K, et al. Gene cloning, expression, and characterization of a novel phytase from Dickeya paradisiaca [J]. Applied Biochemistry and Biotechnology, 2009, 157(2): 113-123. doi: 10.1007/s12010-008-8329-6
[86] de ANGELIS M, GALLO G, CORBO M R, et al. Phytase activity in sourdough lactic acid bacteria: Purification and characterization of a phytase from Lactobacillus sanfranciscensis CB1 [J]. International Journal of Food Microbiology, 2003, 87(3): 259-270. doi: 10.1016/S0168-1605(03)00072-2
[87] AL-ASHEH S, DUVNJAK Z. Characteristics of phytase produced by Aspergillus carbonarius NRC 401121 in canola meal [J]. Acta Biotechnologica, 1994, 14(3): 223-233. doi: 10.1002/abio.370140302
[88] XIANG T, LIU Q, DEACON A M, et al. Crystal structure of a heat-resilient phytase from Aspergillus fumigatus, carrying a phosphorylated histidine [J]. Journal of Molecular Biology, 2004, 339(2): 437-445. doi: 10.1016/j.jmb.2004.03.057
[89] RIBEIRO CORRÊA T L, de QUEIROZ M V, de ARAÚJO E F. Cloning, recombinant expression and characterization of a new phytase from Penicillium chrysogenum [J]. Microbiological Research, 2015, 170: 205-212. doi: 10.1016/j.micres.2014.06.005
[90] TSENG Y H, FANG T J, TSENG S M. Isolation and characterization of a novel phytase from Penicillium simplicissimum [J]. Folia Microbiologica, 2000, 45(2): 121-127. doi: 10.1007/BF02817409
[91] LEE J, CHOI Y, LEE P C, et al. Recombinant production of Penicillium oxalicum PJ3 phytase in Pichia pastoris [J]. World Journal of Microbiology and Biotechnology, 2007, 23(3): 443-446. doi: 10.1007/s11274-006-9236-z
[92] QUAN C S, TIAN W J, FAN S D, et al. Purification and properties of a low-molecular-weight phytase from Cladosporium sp. FP-1 [J]. Journal of Bioscience and Bioengineering, 2004, 97(4): 260-266. doi: 10.1016/S1389-1723(04)70201-7
[93] RAMACHANDRAN S, ROOPESH K, NAMPOOTHIRI K M, et al. Mixed substrate fermentation for the production of phytase byRhizopus spp. using oilcakes as substrates [J]. Process Biochemistry, 2005, 40(5): 1749-1754. doi: 10.1016/j.procbio.2004.06.040
[94] JIN U H, CHUN J A, LEE J W, et al. Expression and characterization of extracellular fungal phytase in transformed sesame hairy root cultures [J]. Protein Expression and Purification, 2004, 37(2): 486-492. doi: 10.1016/j.pep.2004.06.020
[95] BOYCE A, WALSH G. Purification and characterisation of an acid phosphatase with phytase activity from Mucor hiemalis Wehmer [J]. Journal of Biotechnology, 2007, 132(1): 82-87. doi: 10.1016/j.jbiotec.2007.08.028
[96] SINGH B, SATYANARAYANA T. Improved phytase production by a thermophilic mould Sporotrichum thermophile in submerged fermentation due to statistical optimization [J]. Bioresource Technology, 2008, 99(4): 824-830. doi: 10.1016/j.biortech.2007.01.007
[97] SHIMIZU M. Purification and characterization of phytase and acid phosphatase produced by Aspergillus oryzae K1 [J]. Bioscience, Biotechnology, and Biochemistry, 1993, 57(8): 1364-1365. doi: 10.1271/bbb.57.1364
[98] QUAN C S, FAN S D, ZHANG L H, et al. Purification and properties of a phytase from Candida krusei WZ-001 [J]. Journal of Bioscience and Bioengineering, 2002, 94(5): 419-425.
[99] HAN Y M, LEI X G. Role of glycosylation in the functional expression of an Aspergillus niger Phytase (phyA) in Pichia pastoris [J]. Archives of Biochemistry and Biophysics, 1999, 364(1): 83-90. doi: 10.1006/abbi.1999.1115
[100] HELLSTRÖM A M, VÁZQUES-JUÁREZ R, SVANBERG U, et al. Biodiversity and phytase capacity of yeasts isolated from Tanzanian togwa [J]. International Journal of Food Microbiology, 2010, 136(3): 352-358. doi: 10.1016/j.ijfoodmicro.2009.10.011
[101] HARALDSSON A K, VEIDE J, ANDLID T, et al. Degradation of phytate by high-phytase Saccharomyces cerevisiae strains during simulated gastrointestinal digestion [J]. Journal of Agricultural and Food Chemistry, 2005, 53(13): 5438-5444. doi: 10.1021/jf0478399
[102] SEGUEILHA L, LAMBRECHTS C, BOZE H, et al. Purification and properties of the phytase from Schwanniomyces castellii [J]. Journal of Fermentation and Bioengineering, 1992, 74(1): 7-11. doi: 10.1016/0922-338X(92)90259-W
[103] LIM B L, YEUNG P, CHENG C W, et al. Distribution and diversity of phytate-mineralizing bacteria [J]. The ISME Journal, 2007, 1(4): 321-330. doi: 10.1038/ismej.2007.40
[104] ROCKY-SALIMI K, HASHEMI M, SAFARI M, et al. A novel phytase characterized by thermostability and high pH tolerance from rice phyllosphere isolated Bacillus subtilis B. S. 46 [J]. Journal of Advanced Research, 2016, 7(3): 381-390. doi: 10.1016/j.jare.2016.02.003
[105] MUKHAMETZIANOVA A D, AKHMETOVA A I, SHARIPOVA M R. Microorganisms as phytase producers [J]. Mikrobiologiia, 2012, 81(3): 291-300.
[106] YAO M Z, ZHANG Y H, LU W L, et al. Phytases: crystal structures, protein engineering and potential biotechnological applications [J]. Journal of Applied Microbiology, 2012, 112(1): 1-14. doi: 10.1111/j.1365-2672.2011.05181.x
[107] BALWANI I, CHAKRAVARTY K, GAUR S. Role of phytase producing microorganisms towards agricultural sustainability [J]. Biocatalysis and Agricultural Biotechnology, 2017, 12: 23-29. doi: 10.1016/j.bcab.2017.08.010
[108] CHEN C R, CONDRON L M, DAVIS M R, et al. Phosphorus dynamics in the rhizosphere of perennial ryegrass (Lolium perenne L. ) and radiata pine (Pinus radiata D. Don. ) [J]. Soil Biology and Biochemistry, 2002, 34(4): 487-499. doi: 10.1016/S0038-0717(01)00207-3
[109] BÜNEMANN E K. Enzyme additions as a tool to assess the potential bioavailability of organically bound nutrients [J]. Soil Biology and Biochemistry, 2008, 40(9): 2116-2129. doi: 10.1016/j.soilbio.2008.03.001
[110] TARAFDAR J C, YADAV R S, MEENA S C. Comparative efficiency of acid phosphatase originated from plant and fungal sources [J]. Journal of Plant Nutrition and Soil Science, 2001, 164(3): 279-282. doi: 10.1002/1522-2624(200106)164:3<279::AID-JPLN279>3.0.CO;2-L
[111] JORQUERA M A, CROWLEY D E, MARSCHNER P, et al. Identification of β-propeller phytase-encoding genes in culturable Paenibacillus and Bacillus spp. from the rhizosphere of pasture plants on volcanic soils [J]. FEMS Microbiology Ecology, 2011, 75(1): 163-172. doi: 10.1111/j.1574-6941.2010.00995.x
[112] BOHN L, MEYER A S, RASMUSSEN S K. Phytate: impact on environment and human nutrition. A challenge for molecular breeding [J]. Journal of Zhejiang University. Science. B, 2008, 9(3): 165-191. doi: 10.1631/jzus.B0710640
[113] QUIQUAMPOIX H, BURNS R G. Interactions between proteins and soil mineral surfaces: Environmental and health consequences [J]. Elements, 2007, 3(6): 401-406. doi: 10.2113/GSELEMENTS.3.6.401
[114] WALLENSTEIN M, ALLISON S D, ERNAKOVICH J, et al. Controls on the temperature sensitivity of soil enzymes: A key driver of in situ enzyme activity rates [J].Soil Enzymology, 2011: 245–258.
[115] FAROUK A E A, GREINER R, HUSSIN A S M. Purification and properties of a phytate-degrading enzyme produced by Enterobacter sakazakii ASUIA279 [J]. Journal of Biotechnology and Biodiversity, 2012, 3(1): 1-9.
[116] ESCOBIN-MOPERA L, OHTANI M, SEKIGUCHI S, et al. Purification and characterization of phytase from Klebsiella pneumoniae 9-3B [J]. Journal of Bioscience and Bioengineering, 2012, 113(5): 562-567. doi: 10.1016/j.jbiosc.2011.12.010
[117] SREEDEVI S, REDDY B N. Purification and biochemical characterization of phytase from newly isolated Bacillus subtilis C43 [J]. Advanced Biotechnology, 2013, 12(8): 52-60.
[118] KONIETZNY U, GREINER R. Molecular and catalytic properties of phytate-degrading enzymes (phytases) [J]. International Journal of Food Science and Technology, 2002, 37(7): 791-812. doi: 10.1046/j.1365-2621.2002.00617.x
[119] EL-TOUKHY N M K, YOUSSEF A S, MIKHAIL M. Isolation, purification and characterization of phytase from Bacillus subtilis MJA [J]. African Journal of Biotechnology, 2013, 12: 2957-2967.
[120] WYSS M, BRUGGER R, KRONENBERGER A, et al. Biochemical characterization of fungal phytases (myo-inositol hexakisphosphate phosphohydrolases): Catalytic properties [J]. Applied and Environmental Microbiology, 1999, 65(2): 367-373. doi: 10.1128/AEM.65.2.367-373.1999
[121] MARLIDA Y, DELFITA R, ADNADI P, et al. Isolation, characterization and production of phytase from endophytic fungus its application for feed [J]. Pakistan Journal of Nutrition, 2010, 9(5): 471-474. doi: 10.3923/pjn.2010.471.474
[122] CRAINE J M, FIERER N, MCLAUCHLAN K K. Widespread coupling between the rate and temperature sensitivity of organic matter decay [J]. Nature Geoscience, 2010, 3(12): 854-857. doi: 10.1038/ngeo1009
[123] KOCH O, TSCHERKO D, KANDELER E. Temperature sensitivity of microbial respiration, nitrogen mineralization, and potential soil enzyme activities in organic alpine soils [J]. Global Biogeochemical Cycles, 2007, 21(4): 1-11.
[124] WALLENSTEIN M D, MCMAHON S K, SCHIMEL J P. Seasonal variation in enzyme activities and temperature sensitivities in Arctic tundra soils [J]. Global Change Biology, 2009, 15(7): 1631-1639. doi: 10.1111/j.1365-2486.2008.01819.x
[125] LONHIENNE T, GERDAY C, FELLER G. Psychrophilic enzymes: Revisiting the thermodynamic parameters of activation may explain local flexibility [J]. Biochimica et Biophysica Acta (BBA) - Protein Structure and Molecular Enzymology, 2000, 1543(1): 1-10. doi: 10.1016/S0167-4838(00)00210-7
[126] FELLER G. Molecular adaptations to cold in psychrophilic enzymes [J]. Cellular and Molecular Life Sciences:CMLS, 2003, 60(4): 648-662. doi: 10.1007/s00018-003-2155-3
[127] JOHNSON S C, YANG M M, MURTHY P P N. Heterologous expression and functional characterization of a plant alkaline phytase in Pichia pastoris [J]. Protein Expression and Purification, 2010, 74(2): 196-203. doi: 10.1016/j.pep.2010.07.003
[128] KONIETZNY U, GREINER R. Bacterial phytase: Potential application, in vivo function and regulation of its synthesis [J]. Brazilian Journal of Microbiology, 2004, 35(1/2): 12-18.
[129] GEORGE T S, RICHARDSON A E, SIMPSON R J. Behaviour of plant-derived extracellular phytase upon addition to soil [J]. Soil Biology and Biochemistry, 2005, 37(5): 977-988. doi: 10.1016/j.soilbio.2004.10.016
[130] BOGAR B, SZAKACS G, LINDEN J C, et al. Optimization of phytase production by solid substrate fermentation [J]. Journal of Industrial Microbiology and Biotechnology, 2003, 30(3): 183-189. doi: 10.1007/s10295-003-0027-3
[131] TANG J, LEUNG A, LEUNG C, et al. Hydrolysis of precipitated phytate by three distinct families of phytases [J]. Soil Biology and Biochemistry, 2006, 38(6): 1316-1324. doi: 10.1016/j.soilbio.2005.08.021
[132] ADAMS M A, PATE J S. Availability of organic and inorganic forms of phosphorus to lupins (Lupinus spp. ) [J]. Plant and Soil, 1992, 145(1): 107-113. doi: 10.1007/BF00009546
[133] RAO M A, VIOLANTE A, GIANFREDA L. Interaction of acid phosphatase with clays, organic molecules and organo-mineral complexes: Kinetics and stability [J]. Soil Biology and Biochemistry, 2000, 32(7): 1007-1014. doi: 10.1016/S0038-0717(00)00010-9